If we approach territoriality from the perspective of biology, we can use the understanding of territory advanced by the ethologist Jakob von Uexküll. Von Uexküll proposed that there is no meaning outside of a milieu (Umwelt). For him a ‘territory’ refers to a specific milieu that cannot be separated from the living thing occupying and creating the milieu, so that the meaning of a milieu for Von Uexküll is affective.